Macrosemiidae
Macrosemiidae is an extinct family of ginglymodian fish. The family first appeared during the Middle Triassic and disappeared during the Late Cretaceous. The group has been found in rock formations in Africa, Australia, Eurasia and North America. As ginglymodians, their closest living relatives are gars, with them being typically placed as nested within the Semionotiformes. Many members of the family have elongated dorsal fins, often associated with adjacent area of skin which was free of scales. These fins were likely undulated for use in precision swimming. Their body morphology suggests that they were slow swimmers that were capable of maneuvering around complex topography, such as reef environments.

Arratia, G. and Schultze, H.-P. · CC BY-SA 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- MacrosemiiformesPBDB
- Motility
- actively mobilePBDB, for Osteichthyes
Time the record covers
At least 172 million years on record.
The record covers at least 244.4 Mya to 72.2 Mya. No occurrence read is dated outside 247 Mya to 66 Mya.
86 occurrences of Macrosemiidae on record, most of them in the Middle Triassic. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
86 occurrences- Middle Triassic41%
- Late Triassic12%
- Middle Jurassic1%
- Late Jurassic19%
- Early Cretaceous21%
- Late Cretaceous7%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
25 formations
- Meride Formation9
- Solnhofen Formation7
- Cassina Formation5
- Tlayúa Formation4
- Hauptdolomit Formation4
- Vossenveld Formation4
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The named rock units Macrosemiidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. One further formation holds fewer occurrences and is not listed. 27 occurrences have no formation on record, which is common for older literature.
Macrosemiidae and its relatives
Middle TriassicWhere Macrosemiidae and its relatives were recorded in the Middle Triassic.
Found in the same rock
696 taxa- Eosemionotus14 sites
- Legnonotus krambergeri9 sites
- Saurichthys9 sites
- Caturus8 sites
- Eosemionotus diskosomus8 sites
- Pholidophorus8 sites
- Belonostomus6 sites
- Colobodus5 sites
- Paralepidotus ornatus5 sites
- Pholidophorus latiusculus5 sites
- Actinopterygii4 sites
- Archaeosemionotus4 sites
- Brachyphyllum4 sites
- Dandya ovalis4 sites
- Lepidotes4 sites
- Luganoia lepidosteoides4 sites
- Notagogus novomundi4 sites
- Ophiopsiella attenuata4 sites
- Perledovatus nothosomoides4 sites
- Perleidus altolepis4 sites
- Pleuropholis4 sites
- Ptycholepis barboi4 sites
- Pycnodontiformes4 sites
- Acanthomorphata3 sites
The Paleobiology Database records 71 sites for Macrosemiidae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
34 references- Taxonomy and phylogeny of Eosemionotus Stolley, 1920 (Neopterygii: Ginglymodi) from the Middle Triassic of Europe13
- Catalogue of the Fossil Fishes in the British Museum (Natural History), Part III9
- Monographie der fossilien Fische aus den Lithographischen Schiefern Bayerns6
- A new species of Eosemionotus (Holostei, Macrosemiidae) from the Anisian (Middle Triassic) of Winterswijk, the Netherlands4
- The Norian fish deposits of Wiestal ("Seefeld Member", Northern Calcareous Alps, Salzburg, Austria) - taxonomy and palaeoenvironmental implications4
- New holostean fishes (Actinopterygii: Neopterygii) from the Middle Triassic of the Monte San Giorgio (Canton Ticino, Switzerland)4
- An overview of the Middle Triassic actinopterygians from Alcover, Mont-ral and El Pinetell (Catalonia, Spain)4
- The Macrosemiidae (Pisces, Actinopterygii) from the Upper Triassic of Lombardy (N. Italy)4
- A new Notagogus (Macrosemiidae, Halecostomi) species from the Albian Tlayúa quarry, central Mexico3
- Costa, 18643
- Late Cretaceous (Maastrichtian) chondrichthyes and osteichthyes from northeastern Iberia2
- Vertebrate fossils from the Kimmeridgian of Brunn: the oldest fauna from the Solnhofen Archipelago (Late Jurassic, Bavaria, Germany)2
Each of the 86 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 22 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Macrosemiidae
- Source
- Wikipedia/Commons
- Credit
- Arratia, G. and Schultze, H.-P.
- Licence
- CC BY-SA 3.0
- Commercial use
- the licence allows it
No 3D scan of Macrosemiidae was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itMacrosemiiformes through timeFormations of the Middle TriassicThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Thiollière 1858