Pleuronectiformes
Flatfish are a group of ray-finned fish belonging to the suborder Pleuronectoidei of the order Carangiformes. Their collective common name is due to their habit of lying on one side of their laterally-compressed body upon the seafloor; in this position, both eyes lie on the side of the head facing upwards, while the other side of the head and body lies on the substrate. This loss of symmetry, a unique adaptation in vertebrates, stems from one eye "migrating" towards the other during the juvenile's metamorphosis. Due to interspecific variation, some species tend to face their left side upward, some their right side, and others face either side upward; members of Pleuronectidae lie on their left side, with eyes on the right, Paralichthyidae lie on their right side, with eyes on the left, while the "primitive" genus Psettodes may develop into "right-facing" or "left-facing" individuals.

Liné1 · CC BY 2.5 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- flatfishPBDB
- Sits inside
- CarangariaPBDB
- Diet
- carnivorePBDB
- Life habit
- nektobenthicPBDB
- Motility
- actively mobilePBDB
Time the record covers
At least 121 million years on record.
The record covers at least 121.4 Mya to 11.7 ka. No occurrence read is dated outside 125.77 Mya to 0 Mya.
473 occurrences of Pleuronectiformes on record, most of them in the Miocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
473 occurrences- Early Cretaceousunder 1%
- Eocene11%
- Oligocene10%
- Miocene45%
- Pliocene13%
- Pleistocene18%
- Holocene2%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
85 formations
- Palos Verdes Sand Formation20
- Raz-el-Ain Formation13
- Navidad Formation11
- Gatun Formation10
- Port Orford Formation9
- Rifle Butts Formation · Kaierero Group7
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The named rock units Pleuronectiformes has been recovered from, largest first. Drawn where the rock is now, not where it formed. 61 further formations hold fewer occurrences each and are not listed. 228 occurrences have no formation on record, which is common for older literature.
Pleuronectiformes and its relatives
MioceneWhere Pleuronectiformes and its relatives were recorded in the Miocene.
Found in the same rock
4,222 taxa- Diaphus22 sites
- Bregmaceros21 sites
- Apogon18 sites
- Parapercis finlayi18 sites
- Teleostei17 sites
- Pterothrissus umbonatus16 sites
- Hemerocoetes minutus15 sites
- Maurolicus muelleri15 sites
- Acropoma serrata14 sites
- Carcharhinus14 sites
- Clupeidae14 sites
- Conger14 sites
- Gnathophis ornatus14 sites
- Lesueurigobius14 sites
- Merluccius merluccius14 sites
- Raja14 sites
- Arnoglossus13 sites
- Auchenoceros bicaudatus13 sites
- Citharichthys13 sites
- Diaphus splendidus13 sites
- Trachinus13 sites
- Trachurus13 sites
- Cymatogaster aggregata12 sites
- Gobiidae12 sites
The Paleobiology Database records 229 sites for Pleuronectiformes, and this reads all of them. Of that, one read hit the database's own return limit, so every count below is a floor rather than a total. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
133 references- Reconstruction of the fossil marine bony fish fauna (Teleostei) from the Eocene to Pleistocene of New Zealand by means of otoliths. With studies of Recent congroid, morid and trachinoid otoliths60
- Fish otoliths from the early Miocene of Chile: a window into the evolution of marine bony fishes in the Southeast Pacific19
- Otolithes de poissons du paléocanyon de saubrigues (chattien à langhien), aquitaine méridionale, France19
- Fish remains, mostly otoliths and teeth, from the Palos Verdes Sand (Late Pleistocene) of California19
- Données récentes sur Les téléostéens du Miocène et du Pliocéne d'Italie15
- Faunal Horizons of the Barton Beds in Hampshire11
- The Zanclean marine fish fauna and palaeoenvironmental reconstruction of a coastal marine setting in the eastern Mediterranean10
- The Marine Fish Fauna of the Middle Pleistocene Port Orford Formation10
- Messinian marine fish communities of the Mediterranean Sea10
- Fish otoliths from the New Zealand Tertiary10
- Les otolithes des téléostéens de l'oligo-miocène Belge10
- Middle and late Eocene fish otoliths from the eastern and southern USA8
Each of the 473 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 121 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Pleuronectiformes
- Source
- Wikipedia/Commons
- Credit
- Liné1
- Licence
- CC BY 2.5
- Commercial use
- the licence allows it
No 3D scan of Pleuronectiformes was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Flatfish”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itCarangaria through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0.