Xiphosura
Xiphosura is an order of arthropods related to arachnids. They are more commonly known as horseshoe crabs. They first appear in the fossil record in the Early Ordovician, around 480 million years ago. Currently, there are only four living species. Xiphosura contains one suborder, Xiphosurida, and several stem-genera.

Junnn11 · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- ProsomapodaPBDB
- Diet
- carnivorePBDB
- Life habit
- nektobenthicPBDB
- Motility
- actively mobilePBDB
Time the record covers
At least 502 million years on record.
The record covers at least 506.5 Mya to 3.6 Mya. No occurrence read is dated outside 538.8 Mya to 2.58 Mya.
254 occurrences of Xiphosura on record, most of them in the Carboniferous. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
254 occurrences- Cambrianunder 1%
- Ordovician3%
- Silurian4%
- Devonian6%
- Carboniferous57%
- Permian6%
- Early Triassic4%
- Middle Triassic8%
- Late Triassic2%
- Early Jurassic1%
- Middle Jurassicunder 1%
- Late Jurassic6%
- Early Cretaceousunder 1%
- Late Cretaceous2%
- Eoceneunder 1%
- Plioceneunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
106 formations
- Carbondale Formation · Kewanee Group9
- Solnhofen Formation9
- Grès à Voltzia Formation7
- Wellington Formation5
- Wettin Formation5
- Pennant Sandstone Formation4
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The named rock units Xiphosura has been recovered from, largest first. Drawn where the rock is now, not where it formed. 82 further formations hold fewer occurrences each and are not listed. 88 occurrences have no formation on record, which is common for older literature.
Xiphosura and its relatives
CarboniferousWhere Xiphosura and its relatives were recorded in the Carboniferous.
Found in the same rock
1,617 taxa- Euproops danae28 sites
- Belinurus trilobitoides27 sites
- Liomesaspis laevis12 sites
- Xiphosurida10 sites
- Limulitella bronnii9 sites
- Prestwichianella rotundata9 sites
- Belinurus7 sites
- Euproops7 sites
- Mesolimulus walchi6 sites
- Paleolimulus signatus6 sites
- Belinurus iswarinensis5 sites
- Limulidae5 sites
- Palaeocampa anthrax5 sites
- Conulariida4 sites
- Diplostraca4 sites
- Eoscorpiidae4 sites
- Euestheria minuta4 sites
- Euthycarcinus kessleri4 sites
- Insecta4 sites
- Lingularia tenuissima4 sites
- Palaeoxyris regularis4 sites
- Actinopterygii3 sites
- Animalia3 sites
- Antrimpos atavus3 sites
The Paleobiology Database records 216 sites for Xiphosura, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
137 references- On the fossil Xiphosura from the South Wales Coalfield with a note on the myriapod Euphoberia28
- Late Paleozoic xiphosurans12
- The arthropod fauna of the Upper Carboniferous rocks of the Maritime Provinces9
- Some coal measures arthropods from the south Wales coalfield8
- Zametka o predstavitelyakh Xiphosura iz Donetskogo Basseyna7
- Planche VII. Limules fossiles des couches supérieures du Jura7
- Habitat of Llandoverian-Lochkovian eurypterids5
- Arthropodenfunde im Stefan der Halleschen Mulde5
- New data on Carboniferous xiphosurans (Xiphosura, Chelicerata) of the Donets Coal Basin4
- Limulines of the Mississippian Bear Gulch Limestone of Central Montana, USA4
- Faunes et paysages du Grès à Voltzia du Nord des Vosges: Essai paléoécologique sur le Buntsandstein supérieur4
- Eshche o Phyllopoda i Xiphosura Donetskogo basseyna4
Each of the 254 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 125 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Xiphosura
- Source
- Wikipedia/Commons
- Credit
- Junnn11
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Xiphosura was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Xiphosura”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itProsomapoda through timeFormations of the CarboniferousThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Latreille 1802)