Pluvialis squatarola
The grey plover or black-bellied plover is a large plover breeding in Arctic regions. It is a long-distance migrant, with a nearly worldwide coastal distribution when not breeding.

Chuck Homler, Focus On Wildlife · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- black-bellied ploverPBDB
- Sits inside
- PluvialisPBDB
- Diet
- carnivorePBDB, for Coelurosauria
- Life habit
- ground dwelling, solitaryPBDB, for Theropoda
- Motility
- actively mobilePBDB, for Theropoda
- Environment
- marinePBDB, for Charadriiformes
Time the record covers
At least 3.5 million years on record.
The record covers at least 3.6 Mya to 11.7 ka. No occurrence read is dated outside 5.333 Mya to 0 Mya.
17 occurrences of Pluvialis squatarola on record, most of them in the Pleistocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
17 occurrences- Pliocene6%
- Pleistocene65%
- Holocene29%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
1 formation
- Yorktown Formation · Chesapeake Group1
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The named rock units Pluvialis squatarola has been recovered from, largest first. Drawn where the rock is now, not where it formed. 16 occurrences have no formation on record, which is common for older literature.
Pluvialis squatarola and its relatives
PleistoceneWhere Pluvialis squatarola and its relatives were recorded in the Pleistocene.
Found in the same rock
811 taxa- Tyto alba11 sites
- Buteo9 sites
- Corvus corax9 sites
- Haliaeetus leucocephalus9 sites
- Larus9 sites
- Antigone canadensis8 sites
- Buteo jamaicensis8 sites
- Corvus brachyrhynchos8 sites
- Tringa semipalmata8 sites
- Anser caerulescens7 sites
- Athene cunicularia7 sites
- Branta canadensis7 sites
- Limosa fedoa7 sites
- Numenius americanus7 sites
- Phalacrocorax penicillatus7 sites
- Podiceps nigricollis7 sites
- Anas6 sites
- Charadrius vociferus6 sites
- Falco peregrinus6 sites
- Melanitta6 sites
- Melanitta perspicillata6 sites
- Mergus serrator6 sites
- Phalacrocorax6 sites
- Phalacrocorax auritus6 sites
The Paleobiology Database records 17 sites for Pluvialis squatarola, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
11 references- Prehistoric human impacts on California birds: evidence from the Emeryville shellmoumd avifauna5
- Humans and marine resource interaction reappraised: Archaeofauna remains during the late Pleistocene and Holocene in Cantabrian Spain2
- Distribution and provenance of fossil avifauna on San Miguel Island2
- Terminal Pleistocene–Holocene Avifauna of San Miguel and Santa Rosa Islands: Identifications of Previously Unidentified Avian Remains Recovered from Fossil Sites and Prehistoric Cave Deposits1
- Pleistocene fossil vertebrate sites of the South East region of South Australia II1
- Miocene and Pliocene birds from the Lee Creek Mine, North Carolina1
- The non-passerine Pleistocene avifauna of the Talara Tar Seeps, northwestern Peru1
- The late Pleistocene avifauna of La Carolina, southwestern Ecuador1
- A comparison of avian assemblages from individual pits at Rancho La Brea, California1
- The fossil Birds of California1
- The avifauna associated with human remains at Rancho la Brea, California1
Each of the 17 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content2
Picture of Pluvialis squatarola
- Source
- Wikipedia/Commons
- Credit
- Chuck Homler, Focus On Wildlife
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
Structured light scan
- Source
- MorphoSource
- Credit
- LACM Ornithology
- Licence
- In Copyright
- Commercial use
- cannot be read from the licence given — check the record
- Specimen
- Pluvialis squatarola · LACM:Birds:87074
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Grey plover”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPluvialis through timeFormations of the PleistoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Linnaeus 1758)