Psammosteidae
Psammosteidae is an extinct family of flattened, benthic heterostracan vertebrates that lived in marine and estuary environments in Europe, Russia & North America. They arose during the Early Devonian, with the first known genus, Drepanaspis from the Hunsrück lagerstätte. The Psammosteids were the only heterostracans that survived the Upper Frasnian extinction event during the Late Devonian, dying out in the extinction event at the very end of the Devonian.

The original uploader was Apokryltaros at English Wikipedia . · CC BY 2.5 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- PteraspidoideiPBDB
- Motility
- actively mobilePBDB, for Chordata
Time the record covers
At least 87 million years on record.
The record covers at least 410.62 Mya to 323.4 Mya. No occurrence read is dated outside 413.02 Mya to 298.9 Mya.
105 occurrences of Psammosteidae on record, most of them in the Devonian. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
105 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
29 formations
- Aruküla Formation7
- Burtnieki Formation6
- Aruküla Formation6
- Yam-Tesovo Formation5
- Vstrechnaya Formation4
- Gauja Formation4
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The named rock units Psammosteidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 5 further formations hold fewer occurrences each and are not listed. 37 occurrences have no formation on record, which is common for older literature.
Psammosteidae and its relatives
DevonianWhere Psammosteidae and its relatives were recorded in the Devonian.
Found in the same rock
352 taxa- Glyptolepis15 sites
- Psammosteus9 sites
- Asterolepis8 sites
- Cheiracanthus brevicostatus8 sites
- Cheirolepis8 sites
- Psammolepis8 sites
- Acanthodii7 sites
- Pycnosteus7 sites
- Sarcopterygii7 sites
- Tartuosteus7 sites
- Acanthodes6 sites
- Actinolepis tuberculata6 sites
- Actinopterygii6 sites
- Byssacanthus dilatatus6 sites
- Cheiracanthus6 sites
- Diplacanthus6 sites
- Homostius latus6 sites
- Orvikuina vardiaensis6 sites
- Asterolepis estonica5 sites
- Devononchus concinnus5 sites
- Ganosteus artus5 sites
- Onychodus5 sites
- Psammolepis toriensis5 sites
- Tartuosteus giganteus5 sites
The Paleobiology Database records 56 sites for Psammosteidae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
26 references- The Middle Devonian fishes of the Baltic States (Estonia, Latvia) and Belarus28
- Upper Silurian and Devonian heterostracan pteraspidomorphs (Vertebrata) from Severnaya Zemlya (Russia): a preliminary report with biogeographical and biostratigraphical implications11
- A new species of the Devonian actinopterygian fish Moythomasia from Belarus10
- Devonian psammosteids (Agnatha, Psammosteidae) of the USSR8
- New tetrapodomorph vertebrates from the Yam-Tesovo locality (Amata Regional Stage, Middle–Upper Devonian) of Leningrad Region, northwestern Russia5
- A new psammosteid (Agnatha, Heterostraci) from the Amata Regional Stage of the Main Devonian Field and morpho-histological types of discrete micromeric elements in the family Psammosteidae5
- Vertebrata, in The Development and Change of Marine Organisms at the Palaeozoic-Mesozoic Boundary5
- Psammosteiformes (Agnatha). A review with descriptions of new material from the Lower Devonian of Poland5
- Late Devonian fossil fishes sites of Scotland4
- New Early Devonian (late Emsian) placoderms from Belarus3
- A new species of ray-finned fish (Osteichthyes, Actinopterygii) from the upper Emsian and lowermost Eifelian deposits of Belarus2
- The Givetian vertebrate fauna from the Fiskekløfta Member (Mimerdalen Subgroup), Svalbard2
Each of the 105 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 14 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Psammosteidae
- Source
- Wikipedia/Commons
- Credit
- The original uploader was Apokryltaros at English Wikipedia .
- Licence
- CC BY 2.5
- Commercial use
- the licence allows it
No 3D scan was found. Smithsonian Open Access, MorphoSource and Sketchfab were all searched.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itPteraspidoidei through timeFormations of the DevonianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Traquair 1896