Pteraspidiformes
Pteraspidiformes is an extinct order of heterostracan agnathan vertebrates known from extensive fossil remains primarily from Early Devonian strata of Europe and North America, and from Upper Silurian Canada.

Apokryltaros · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- HeterostraciPBDB
- Motility
- actively mobilePBDB, for Chordata
Time the record covers
At least 99 million years on record.
The record covers at least 422.7 Mya to 323.4 Mya. No occurrence read is dated outside 432.9 Mya to 298.9 Mya.
349 occurrences of Pteraspidiformes on record, most of them in the Devonian. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
349 occurrences- Silurian2%
- Devonian97%
- Carboniferousunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
73 formations
- Dniester Formation34
- Tyver Formation19
- Psammosteus Limestone Formation18
- Beartooth Butte Formation15
- Wood Bay Formation12
- Ben Nevis Formation11
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The named rock units Pteraspidiformes has been recovered from, largest first. Drawn where the rock is now, not where it formed. 49 further formations hold fewer occurrences each and are not listed. 58 occurrences have no formation on record, which is common for older literature.
Pteraspidiformes and its relatives
DevonianWhere Pteraspidiformes and its relatives were recorded in the Devonian.
Found in the same rock
924 taxa- Protopteraspis gosseleti19 sites
- Cephalaspis18 sites
- Acanthodii16 sites
- Corvaspis15 sites
- Glyptolepis15 sites
- Traquairaspis15 sites
- Turinia pagei12 sites
- Nikolivia elongata10 sites
- Poraspis10 sites
- Protopteraspis10 sites
- Actinopterygii9 sites
- Cheiracanthus brevicostatus9 sites
- Psammosteus9 sites
- Acanthodes8 sites
- Asterolepis8 sites
- Cheirolepis8 sites
- Phialaspis symondsi8 sites
- Podolaspis lerichei8 sites
- Psammolepis8 sites
- Pteraspididae8 sites
- Zascinaspis heintzi8 sites
- Brachipteraspis latissima7 sites
- Byssacanthus dilatatus7 sites
- Cheiracanthus7 sites
The Paleobiology Database records 187 sites for Pteraspidiformes, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
72 references- The Early Devonian armoured agnathans of Podolia, Ukraine57
- The Middle Devonian fishes of the Baltic States (Estonia, Latvia) and Belarus28
- Upper Silurian and Devonian heterostracan pteraspidomorphs (Vertebrata) from Severnaya Zemlya (Russia): a preliminary report with biogeographical and biostratigraphical implications22
- Protopteraspis Gosseleti (Vertebrata: Pteraspidomorphi: Heterostraci) from the Lower Devonian of Shropshire, England17
- Review of the Pteraspidiform heterostracans (Vertebrata, Agnatha) from the Devonian of Podolia, Ukraine, in the Theodor Vascautanu Collection, Bucharest, Romania16
- The vertebrate stratigraphy of the Lower Devonian (Red Bay Group and Wood Bay Formation) of Spitsbergen15
- A new species of the Devonian actinopterygian fish Moythomasia from Belarus11
- A Fossil Flora from the Beartooth Butte Formation of Northern Wyoming9
- Chapter 4. Early Devonian fossil fishes sites of the Welsh Borders8
- Devonian psammosteids (Agnatha, Psammosteidae) of the USSR8
- Habitat of Llandoverian-Lochkovian eurypterids7
- The Osteostraci (Vertebrate, Agnatha) from the Lower Devonian of the Rhenish Slate Mountains, with special reference to their anatomy and phylogenetic position7
Each of the 349 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 60 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Pteraspidiformes
- Source
- Wikipedia/Commons
- Credit
- Apokryltaros
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Pteraspidiformes was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itHeterostraci through timeFormations of the DevonianThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Berg 1940