Spheniscidae
Penguins are a group of flightless semi-aquatic sea birds which live almost exclusively in the Southern Hemisphere. Only one species, the Galapagos penguin, lives at, and slightly north of, the equator. Highly adapted for life in the ocean water, penguins have countershaded dark and white plumage and flippers for swimming. Most penguins feed on krill, fish, squid and other forms of sea life which they catch with their bills and swallow whole while swimming. A penguin has a spiny tongue and powerful jaws to grip slippery prey.

Godot13 · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- SphenisciformesPBDB
- Diet
- carnivorePBDB, for Coelurosauria
- Life habit
- ground dwelling, solitaryPBDB, for Theropoda
- Motility
- actively mobilePBDB, for Theropoda
- Environment
- marinePBDB, for Sphenisciformes
Time the record covers
At least 48 million years on record.
The record covers at least 48.07 Mya to 11.7 ka. No occurrence read is dated outside 56 Mya to 0 Mya.
191 occurrences of Spheniscidae on record, most of them in the Miocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
191 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
38 formations
- La Meseta Formation18
- Pisco Formation14
- Varswater Formation12
- Submeseta Formation12
- Bahía Inglesa Formation11
- Hillgrove Formation8
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The named rock units Spheniscidae has been recovered from, largest first. Drawn where the rock is now, not where it formed. 14 further formations hold fewer occurrences each and are not listed. 57 occurrences have no formation on record, which is common for older literature.
Spheniscidae and its relatives
MioceneWhere Spheniscidae and its relatives were recorded in the Miocene.
Found in the same rock
1,095 taxa- Phalacrocorax16 sites
- Myliobatis13 sites
- Carcharodon hastalis12 sites
- Otodus megalodon12 sites
- Ardenna grisea11 sites
- Larus dominicanus11 sites
- Palaeeudyptes11 sites
- Phocidae11 sites
- Squatina11 sites
- Eudyptula minor10 sites
- Pelagornithidae10 sites
- Pelecanoides urinatrix10 sites
- Spheniscus10 sites
- Cetotheriidae9 sites
- Pachyptila turtur9 sites
- Palaeeudyptes gunnari9 sites
- Pristiophorus9 sites
- Sula9 sites
- Carcharodon carcharias8 sites
- Cetacea8 sites
- Eudyptes pachyrhynchus8 sites
- Isurus8 sites
- Monachinae8 sites
- Anas chlorotis7 sites
The Paleobiology Database records 121 sites for Spheniscidae, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
96 references- Late-Pleistocene avifaunas from Cape Wanbrow, Otago, South Island, New Zealand9
- Annotated catalogue of the fossil vertebrates from Antarctica housed in the Museo de la Plata, Argentina. I. Birds and Land Mammals from La Meseta Formation (Eocene - Early Oligocene)9
- Fossil Penguins9
- A review of the pre-Pliocene penguins of New Zealand7
- A review of the Tertiary fossil Cetacea (Mammalia) localities in Australia6
- A remarkable fossil and archaeological avifauna from Marfells Beach, Lake Grassmere, South Island, New Zealand6
- The evolution of seabirds in the Humboldt Current: new clues from the Pliocene of Central Chile5
- Holocene fossil bird remains from subantarctic Macquarie Island5
- Mid-Holocene macrofossil-bearing raised marine beaches at Potter Peninsula, King George Island, South Shetland Islands5
- Taxonomic revision of Eocene Antarctic penguins based on tarsometatarsal morphology5
- Eocene birds from the western margin of southernmost South America4
- Relict or colonizer? Extinction and range expansion of penguins in southern New Zealand3
Each of the 191 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 84 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Spheniscidae
- Source
- Wikipedia/Commons
- Credit
- Godot13
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Spheniscidae was found in MorphoSource and Sketchfab.
Smithsonian Open Access was not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Penguin”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itSphenisciformes through timeFormations of the MioceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. Bonaparte 1831