Sphenisciformes
Penguins are a group of flightless semi-aquatic sea birds which live almost exclusively in the Southern Hemisphere. Only one species, the Galapagos penguin, lives at, and slightly north of, the equator. Highly adapted for life in the ocean water, penguins have countershaded dark and white plumage and flippers for swimming. Most penguins feed on krill, fish, squid and other forms of sea life which they catch with their bills and swallow whole while swimming. A penguin has a spiny tongue and powerful jaws to grip slippery prey.

Godot13 · CC BY-SA 4.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Also called
- penguinPBDB
- Sits inside
- ProcellariimorphaePBDB
- Diet
- carnivorePBDB, for Coelurosauria
- Life habit
- ground dwelling, solitaryPBDB, for Theropoda
- Motility
- actively mobilePBDB, for Theropoda
- Environment
- marinePBDB
Time the record covers
At least 59 million years on record.
The record covers at least 59.24 Mya to 11.7 ka. No occurrence read is dated outside 66 Mya to 0 Mya.
317 occurrences of Sphenisciformes on record, most of them in the Eocene. The Paleobiology Database records this as a living group. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
317 occurrencesThese are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
52 formations
- La Meseta Formation40
- Submeseta Formation31
- Gaiman Formation21
- Varswater Formation17
- Kokoamu Greensand Formation15
- Pisco Formation14
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The named rock units Sphenisciformes has been recovered from, largest first. Drawn where the rock is now, not where it formed. 28 further formations hold fewer occurrences each and are not listed. 66 occurrences have no formation on record, which is common for older literature.
Sphenisciformes and its relatives
EoceneWhere Sphenisciformes and its relatives were recorded in the Eocene.
Found in the same rock
1,254 taxa- Spheniscidae42 sites
- Phalacrocorax16 sites
- Squatina15 sites
- Myliobatis14 sites
- Carcharodon hastalis13 sites
- Otodus megalodon12 sites
- Ardenna grisea11 sites
- Larus dominicanus11 sites
- Palaeeudyptes11 sites
- Pelagornithidae11 sites
- Phocidae11 sites
- Cetacea10 sites
- Chondrichthyes10 sites
- Eudyptula minor10 sites
- Pelecanoides urinatrix10 sites
- Spheniscus10 sites
- Aves9 sites
- Cetotheriidae9 sites
- Pachyptila turtur9 sites
- Palaeeudyptes gunnari9 sites
- Pristiophorus9 sites
- Sula9 sites
- Carcharhinus brachyurus8 sites
- Carcharodon carcharias8 sites
The Paleobiology Database records 173 sites for Sphenisciformes, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
130 references- Taxonomic revision of Eocene Antarctic penguins based on tarsometatarsal morphology18
- Fossil Penguins18
- Annotated catalogue of the fossil vertebrates from Antarctica housed in the Museo de la Plata, Argentina. I. Birds and Land Mammals from La Meseta Formation (Eocene - Early Oligocene)17
- Early Tertiary penguins of New Zealand10
- New fossil penguins (AVes, Sphenisciformes) from the Oligocene of New Zealand reveal the skeletal plan of stem penguins9
- Late-Pleistocene avifaunas from Cape Wanbrow, Otago, South Island, New Zealand9
- A review of the pre-Pliocene penguins of New Zealand8
- Fossil penguins form the mid-Tertiary of Seymour Island8
- Weddellian marine/coastal vertebrates diversity from a basal horizon (Ypresian, Eocene) of the Cucullaea I Allomember, La Meseta formation, Seymour (Marambio) Island, Antarctica6
- A review of the Tertiary fossil Cetacea (Mammalia) localities in Australia6
- A remarkable fossil and archaeological avifauna from Marfells Beach, Lake Grassmere, South Island, New Zealand6
- Largest-known fossil penguin provides insight into the early evolution of sphenisciform body size and flipper anatomy5
Each of the 317 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 118 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Sphenisciformes
- Source
- Wikipedia/Commons
- Credit
- Godot13
- Licence
- CC BY-SA 4.0
- Commercial use
- the licence allows it
No 3D scan of Sphenisciformes was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Source
- Wikipedia, “Penguin”
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itProcellariimorphae through timeFormations of the EoceneThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Sharpe 1891)