Sphenodontia
Rhynchocephalia is an order of lizard-like reptiles that includes only one living species, the tuatara of New Zealand. Despite its current lack of diversity, during the Mesozoic rhynchocephalians were a speciose group with high morphological and ecological diversity. The oldest record of the group is dated to the Middle Triassic around 244 million years ago, and they had achieved global distribution by the Early Jurassic. Most rhynchocephalians belong to the suborder Sphenodontia ('wedge-teeth'). Their closest living relatives are lizards and snakes in the order Squamata, with the two orders being grouped together in the superorder Lepidosauria.

KeresH · CC BY 3.0 · via Wikipedia/Commons. May be a photograph of a specimen, or an artist’s reconstruction — an interpretation of a skeleton, since colour and soft tissue are almost never preserved.
On record
Recorded, not inferred- Sits inside
- LepidosauriaPBDB
- Motility
- actively mobilePBDB, for Osteichthyes
- Environment
- terrestrialPBDB, for Amniota
Time the record covers
At least 155 million years on record.
The record covers at least 227.3 Mya to 72.2 Mya. No occurrence read is dated outside 237 Mya to 66 Mya.
178 occurrences of Sphenodontia on record, most of them in the Late Triassic. The Paleobiology Database records this as a living group, and the chart stops with its fossils rather than at the present. Bar height is expected occurrences, not a count: each fossil is spread across the span it is dated to, so a tall narrow bar means tight dating and a low wide one means loose.
When
178 occurrences- Late Triassic44%
- Early Jurassic18%
- Middle Jurassic3%
- Late Jurassic30%
- Early Cretaceous5%
- Late Cretaceousunder 1%
These are dates about fossils, not the animal: the record can only begin after it arose and end before it died out, so either end may reach further. Shares are expected occurrences: a fossil dated to a span is counted partly in each period it straddles. More on what this infers.
Found in
36 formations
- Morrison Formation26
- La Boca Formation · Huizachal Group10
- Solnhofen Formation8
- McCoy Brook Formation · Meriden Group6
- Tecovas Formation · Chinle Group4
- Ksar Metlili Formation4
Loading coastlines…
The named rock units Sphenodontia has been recovered from, largest first. Drawn where the rock is now, not where it formed. 12 further formations hold fewer occurrences each and are not listed. 63 occurrences have no formation on record, which is common for older literature.
Sphenodontia and its relatives
Late TriassicWhere Sphenodontia and its relatives were recorded in the Late Triassic.
Found in the same rock
997 taxa- Theropoda27 sites
- Opisthias25 sites
- Clevosaurus21 sites
- Mammalia20 sites
- Actinopterygii17 sites
- Goniopholis17 sites
- Pterosauria17 sites
- Anura15 sites
- Camarasaurus13 sites
- Archosauria12 sites
- Caudata12 sites
- Crocodylia12 sites
- Diphydontosaurus12 sites
- Lepidosauria12 sites
- Ornithischia12 sites
- Testudines12 sites
- Glyptops10 sites
- Kuehneosaurus10 sites
- Ornithopoda10 sites
- Pleurosaurus goldfussi10 sites
- Squamata10 sites
- Allosaurus9 sites
- Crocodylomorpha9 sites
- Dryolestidae9 sites
The Paleobiology Database records 122 sites for Sphenodontia, and this reads all of them. The number is how many of those sites it was catalogued from too. The 24 sharing the most are listed.
This is not a list of everything alive at the time. It is only what has come out of the same ground as this animal and been written down.
Entered from
87 references- Assemblages of small tetrapods from British Late Triassic fissure deposits17
- The ecology of a Late Triassic reptile assemblage from Gloucestershire, England11
- Mesozoic lepidosauromorphs of Mexico: a review and discussion of taxonomic assignments10
- Paleoecological analysis of the vertebrate fauna of the Morrison Formation (Upper Jurassic), Rocky Mountain region, U.S.A.10
- Les rhynchocephales et les pterosauriens a crete parietale du Kimeridgien superieur-Berriasien d'Europe occidentale: le gisement de Canjuers (Var-France) et ses abords8
- Les Rhynchocéphales et les Sauriens des Calcaires lithographiques (Jurassique supérieur) d’Europe occidentale6
- A distinctive Late Triassic microvertebrate fissure fauna and a new species of Clevosaurus (Lepidosauria: Rhynchocephalia) from Woodleaze Quarry, Gloucestershire, UK4
- Late Triassic microvertebrates from the lower Chinle Group (Otischalkian-Adamanian: Carnian), southwestern U.S.A.4
- The Late Triassic lepidosaur fauna from Hallau, North-Eastern Switzerland, and a new ‘basal’ rhynchocephalian Deltadectes elvetica gen. et sp. nov.3
- The age, fauna and palaeoenvironment of the Late Triassic fissure deposits of Tytherington, South Gloucesterhire, UK3
- Connely, 20023
- Marine members of the Sphenodontia3
Each of the 178 occurrences read was entered from one of these publications, ordered by how many each accounts for. This is not a search for papers mentioning the name. 75 further references are not listed. Entries link to doi.org where the Paleobiology Database holds an identifier, and to its own record where it does not. Relayed as the Paleobiology Database holds them, not checked against what they resolve to.
Media and terms
Media content1
Picture of Sphenodontia
- Source
- Wikipedia/Commons
- Credit
- KeresH
- Licence
- CC BY 3.0
- Commercial use
- the licence allows it
No 3D scan of Sphenodontia was found in Sketchfab.
Smithsonian Open Access and MorphoSource were not reached for this taxon, so this list is what the others held rather than everything there is.
The description on this site
- Licence
- CC BY-SA 4.0
This is what the site found, which is not the same as what exists.
Keep going
What may have been alongside itLepidosauria through timeFormations of the Late TriassicThe world it lived on
Occurrence data from the Paleobiology Database, CC BY 4.0. (Jaekel 1910)